The embryological origins of the gene theory / Scott F. Gilbert.
Abstract:
"The original support for the gene theory of inheritance came largely from the studies of E. B. Wilson, Theodor Boveri, and Thomas Hunt Morgan. Each of these scientists began his career as an embryologist. In this paper, the emergence of the gene theory out of embryology will be seen in the context of these researchers' attempts to solve the problem of which cellular compartment--the nucleus or the cytoplasm--directed development."
Journal of the history of biology T. 11 (1978), p. 307-351